The epaulette shark of Australian reefs walks across exposed coral flats on its paddle-shaped fins, wriggling between tide pools. It survives the trip by tolerating extreme oxygen loss that would injure most vertebrates, shutting down non-essential brain functions until it reaches water. It is a fish that hikes between puddles.
When food is thick, a whale shark can hang nearly vertical in the water and gulp — using suction rather than forward motion to draw in prey-rich water. Snorkelers at feeding aggregations often meet the giants bobbing upright like feeding balloons. The largest fish can dine without swimming an inch.
Megalodon, the extinct giant that grew several times the size of any shark alive, left teeth so like a great white shark's that the family tree looks obvious — but fossil evidence places the two on separate lineages, the white shark's ancestry tracking closer to ancient mako-like sharks. Megalodon's line ended; it left no descendants. The great white is the giant's rival's grandchild, not the giant's.
Dried, untanned shark skin — shagreen, usually from small sharks called dogfish — is a natural abrasive: its denticles cut like 80- to 220-grit paper in one direction and burnish in the other, and cabinetmakers and instrument makers smoothed wood with it for centuries; a microscope study of a 1711 Stradivari cello found what looks like a dogfish denticle lodged in the varnish. Tanned and dyed, shagreen also covered cases, boxes and sword grips. The famous exception is the samurai sword: its hilt covering is called samegawa, “shark skin,” but the museum pieces are wrapped in ray skin. On the katana the shark got the name and the ray did the work.
The deep-sea goblin shark's jaws shoot forward at prey in what researchers measured as the fastest jaw protrusion known in fishes, snapping out from under its blade-like snout before retracting. Video analysis published in 2016 called the mechanism "slingshot feeding." Its face is a loaded weapon.
Jelly-filled pores on a shark's snout — the ampullae of Lorenzini — detect the faint electric fields of living bodies: in Kalmijn's 1982 experiments, sharks and rays responded to voltage gradients of just 5 nanovolts per centimeter, five billionths of a volt. A flounder hiding under sand still broadcasts its heartbeat. To a shark, every animal in the sea is a battery that cannot be switched off.
Named for its schedule, the night shark spends days hundreds of meters down and rises toward the surface after dark, guided by oversized eyes with a vivid green shine. Fishermen named it before scientists explained it. Its workday begins when the ocean's lights go out.
Three-dimensional tracking of octopus-fish hunting groups showed goatfish scouting ahead and signaling discoveries, while the octopus largely determined whether the group moved toward them. Group composition also changed how much each species contributed and how successful the hunt was.
Helicoprion, a 270-million-year-old shark relative, grew a spiral whorl of teeth like a circular saw blade set in its lower jaw — a structure so bizarre that scientists argued for a century about where on the body it belonged until CT scans settled it in 2013. Evolution once shipped a power tool with fins.
Unlike whale sharks, which gulp and suction-feed, the basking shark is a pure ram filter feeder: its colossal gape works only when the animal swims forward, forcing water through its gill rakers. Stop the shark and the filter stops with it. For a basking shark, motion is the meal.
Cirrate octopuses, the deep-sea group nicknamed dumbo for the ear-like fins above their eyes, propel themselves by slowly beating those fins rather than by jetting water. A web of skin joins the arms into an umbrella. The deepest octopus ever filmed was one of them, foraging almost 7 kilometers down.
Vertebrate and cephalopod camera eyes evolved independently, but their retinas are wired differently. In vertebrates, nerve fibers lie in front of the photoreceptors and the optic nerve creates a blind spot. In octopuses, the photoreceptors face incoming light and the nerve fibers exit behind them, so there is no comparable blind spot.
Behind a shark's retina lies the tapetum lucidum, a layer of mirror-like plates that bounces unabsorbed light back through the photoreceptors for a second pass. The same structure makes a cat's eyes shine in headlights. In dim water, a shark's eye reads every photon twice.
The most rigorous global estimate, published in Science in 2024, found fishing kills roughly 80 million sharks annually — about 25 million of them from threatened species — and the toll rose even as anti-finning laws increased tenfold. Sharks, by comparison, kill about six people worldwide in a typical year. For every human lost to a shark, humans take more than 10 million sharks.
Expeditions to Kavachi, an active undersea volcano in the Solomon Islands, filmed hammerheads and silky sharks living inside its acidic, superheated crater plume — a discovery nicknamed "Sharkcano" that NASA later observed erupting from orbit. Some sharks chose the one neighborhood everything else evacuated.
In cephalopod anatomy, arms carry suckers along their whole length while tentacles are longer, retractable and suckered mainly at the tip. Squid and cuttlefish have both — eight arms plus two tentacles. Octopuses have eight arms and nothing else. The word "tentacle" is almost universally applied to them and is almost universally wrong.
Venus turns so slowly that a single rotation takes 243 Earth days, while its whole trip around the Sun takes only 225. On Venus, the day outlasts the year.
The animal long called the common octopus, Octopus vulgaris, is now understood as a species complex rather than a single worldwide species. Genetic studies have identified at least six regional species that can be extremely difficult to distinguish by appearance alone.
At the Seattle Aquarium, eight giant Pacific octopuses met two unfamiliar people dressed the same way for two weeks: one always fed them, the other always touched them with a bristly stick. By the end, the animals showed measurably different body patterns, arm movements and funnel orientation toward each person, and the larger ones changed their breathing rate. The uniform trick was the point. They were not reading the clothes.
Fossil sites in Panama, Spain and the southeastern U.S. preserve concentrations of small juvenile teeth of megalodon, the extinct giant shark that hunted the oceans until about 3.6 million years ago — evidence the giants birthed and raised young in warm, food-rich shallows; 2025 growth analyses indicate newborns already stretched about 3.6–3.9 m (12–13 ft). The deadliest daycare in history had ocean views.
When the Moon formed about 4.5 billion years ago, the young Earth spun so fast that a full day lasted only around six hours. The Moon's tides have been braking the planet ever since — for about a billion years in the middle, the day even stalled near 19 hours while sun-driven tides in the atmosphere pushed back. Today the day is still stretching, by about two milliseconds every century.
A shark's entire skeleton is cartilage — the same flexible tissue as a human ear — reinforced with mineralized plates where stress is highest. Cartilage is lighter and bends without breaking, part of why sharks are such economical swimmers. It is also why complete shark fossils are so rare while their teeth litter the world.
An octopus can rebuild a lost arm, including its muscles, suckers and nerve cord. The wound is first covered by a layer of epithelium, then new tissue is organized into the structures of a functional replacement arm rather than a simple scar.
The giant Pacific octopus (Enteroctopus dofleini) can span about 4.3 meters (14 feet) from arm to arm. The heaviest individual reliably weighed alive was 71 kilograms (156 pounds), while most adults are far smaller.
Female sharks of several species can store viable sperm in specialized glands for months to years after mating — one brownbanded bamboo shark produced a pup after 45 months of isolation, a record documented in 2015. Shark family planning operates on geological patience.
With no internal or external skeleton, an octopus is roughly 90% muscle. Its arms act as muscular hydrostats, stiffening, bending and extending by contracting muscles against one another rather than pulling on bones. This structure gives the animal both strength and extreme flexibility.
Tagged blue sharks ride ocean gyres on migrations spanning thousands of miles — Atlantic individuals recaptured across the ocean from where they were marked — following temperature bands and prey like long-haul truckers of the open sea. Their round trips can circle an entire ocean basin.
Satellite-tagged whale sharks travel thousands of kilometers to arrive precisely when food blooms — coral spawn off Australia, fish eggs off Yucatán — forming seasonal gatherings of hundreds, the largest known assemblies of giant fish. Somehow, the world's biggest fish keeps a calendar.
Researchers measuring shark bites found the bull shark delivering nearly 6,000 newtons (about 1,300 pounds of force) at the back teeth — the strongest bite for body size of any shark tested, published in 2012. The stout jaw is built like a bolt cutter for turtles and big fish. Size sets the headline; the bull shark sets the ratio.
Sharks are tuned to low-frequency pulses — the irregular vibrations of a struggling fish — which they detect with inner ears and a lateral-line system running the length of the body. Long before a shark sees you, it has heard and felt you.
Archaeologists have opened Egyptian tombs and found pots of honey more than 3,000 years old — still perfectly edible. With almost no moisture, heavy sugar and a naturally acidic pH, honey is a pantry where bacteria simply cannot live.
Field measurements put the Greenland shark's heart rate near one beat every ten seconds, among the slowest recorded for any fish — machinery throttled down for water near freezing. The same economy drives its glacial growth and centuries-long life. It is an animal living in slow motion.
Off South Africa, a pair of orcas has repeatedly killed great white sharks and removed their energy-rich livers with almost surgical precision, leaving the rest. Tagging studies show white sharks fleeing the region for months after such attacks. The ocean's most feared fish has a boss.
Great white sharks, shortfin makos and their mackerel-shark kin run heat exchangers of tightly packed blood vessels that keep swimming muscles, eyes and brain well above sea temperature — regional warm-bloodedness in a fish. A warm retina sees fast prey in cold water. The great white hunts pre-heated.
Flat as a doormat, the angel shark buries itself in sand with only eyes showing and explodes upward in a tenth-of-a-second strike on passing fish. Once common from Norway to the Canaries and across the Mediterranean, it has disappeared from about 90% of that range in a century, was listed Critically Endangered by the IUCN in 2006, and hangs on mainly around the Canary Islands. The floor of the sea sometimes has teeth.
The eel-like frilled shark, a deep-water relic with 300 needle teeth arranged in frills, is estimated to carry its young for around 3.5 years — among the longest gestations proposed for any vertebrate. In the cold deep, even birth runs in slow motion.
Sharks produce unusually small, rugged antibody fragments called VNARs — roughly a tenth the size of human antibodies — able to reach folds on viral proteins that larger molecules cannot. Lab studies have shown shark VNARs neutralizing SARS-CoV-2 and related coronaviruses, and researchers are developing them against cancers and other targets. The ocean's oldest immune system is auditioning for ours.
Octopuses use copper-based hemocyanin instead of iron-based hemoglobin to carry oxygen. Oxygenated hemocyanin is blue and remains effective in cold, oxygen-poor water, although it carries less oxygen per unit of blood than hemoglobin.
A whale shark's cavernous mouth carries about 300 rows of tiny vestigial teeth — thousands in all — that play no role in feeding, leftovers from toothier ancestors; the animal filter-feeds instead through gill rakers. Evolution rarely cleans out the attic.
The shortfin mako carries its Māori name into science and sport worldwide; in New Zealand tradition mako teeth were treasured ear pendants of high status. One of the ocean's fastest animals is one of the few whose everyday English name is borrowed from a Polynesian language.
CT scans published in 2021 revealed that the spiral intestine of sharks functions like a Tesla valve, easing food one way while resisting backflow — a design engineers patented a century after sharks evolved it. Digestion, in a shark, is plumbing genius.
Biomechanical modeling published in 2008 estimated the bite force of megalodon, the largest shark that ever lived, at roughly 108,000 to 182,000 newtons — up to about 41,000 pounds of force, the strongest bite calculated for any animal at the time. That is several times the estimated bite of Tyrannosaurus rex. The figure often circulates mislabeled as “psi”; it is total force, and it needed no exaggerating.
The siphon is a muscular funnel that forcefully expels water from the body. An octopus uses it for jet propulsion, to eject ink, and to aim water beneath the arm web strongly enough to throw silt, shells and other debris.
In Hawaiian tradition, certain sharks were ʻaumākua — ancestral guardian spirits of specific families — fed, respected and never hunted by those they protected. The man-eater of one culture's nightmares was another culture's grandmother.
Global assessments led by IUCN specialists conclude that roughly one in three species of sharks, rays and chimaeras is threatened with extinction, chiefly from overfishing. The animals that outlasted five mass extinctions are struggling with a sixth made of nets and hooks.
The male seven-arm octopus (Haliphron atlanticus) hides its eighth arm, a specialized mating arm, in a pouch beneath the right eye, so only seven are visible. This deep-sea species feeds on jellyfish, eating the bell and sometimes carrying the trailing tentacles.
Detached arms continue responding to stimuli and activating their suckers long after they lose contact with the brain — in laboratory observations, for more than an hour. In one experiment an octopus handled its own amputated arm freely but held it only at the skinless cut end and would not eat it. The self-recognition signal apparently survives the separation.
In the late 1950s, Eugenie Clark trained lemon sharks at Florida's Cape Haze laboratory to push a target that rang a bell for food, then to pick the correct target over a wrong one — and after a 10-week winter layoff, the sharks took the task straight back up. In 2018, Port Jackson sharks in a Macquarie University lab learned to swim to a feeding station whenever a jazz track played, though asked to tell jazz from classical, they struggled. The reputation for mindless instinct does not survive contact with a curious shark.
Researcher Jennifer Mather observed a wild octopus leave its den, collect three rocks, set them aside, carry them back and stack them across the entrance before resting. The sequence has been discussed as possible planning, although that interpretation remains debated.
Sawsharks wield flattened, tooth-edged snouts hung with a pair of trailing barbels, slashing through fish schools and raking sand for buried prey — unlike the larger sawfish (a ray) they resemble. It is a weapon, a metal detector and a shovel in one nose.
With pectoral fins shaped like wings and a tail delivering forward thrust, many sharks fly through water more than they float in it — negatively buoyant bodies held aloft by hydrodynamics, sinking whenever they glide too slowly. A great white shark is, structurally, a heavier-than-water aircraft.
In a 2021 experiment, 20 wild-caught bonnethead sharks held in a tank were exposed to magnetic fields mimicking spots hundreds of kilometers (hundreds of miles) from where they were caught — and turned to swim “home,” orienting on magnetic cues alone, the first solid evidence that sharks read Earth's magnetic field as a true map. Some of the longest migrations in the sea may run on planetary GPS.
Female bull sharks deliver pups in estuaries and river mouths, and the young may spend years in low-salinity nurseries where big marine predators cannot follow. Fresh water is not a stunt for the species; it is a life stage. Some of the ocean's most formidable sharks learned to swim in rivers.
The basking shark, the second-largest fish, cruises open-mouthed and passively strains plankton from roughly 1,800 metric tons of seawater per hour — on the order of an Olympic pool's worth. All that machinery runs on animals barely visible to the eye.
Before petroleum, oil rendered from shark livers lit lamps, tanned leather and greased machinery; entire fisheries — from Achill Island's basking sharks to the World War II hunt for vitamin A-rich soupfin sharks — rose and crashed on shark oil. Sharks powered human nights before electricity did.
When basking shark carcasses decay, the gill apparatus and lower jaw fall away first, leaving a small-headed, long-necked hulk — the source of several famous "sea serpent" and "plesiosaur" strandings, including the 1977 Zuiyo-maru carcass hauled up off New Zealand. Many monsters, examined closely, turn out to be sharks twice over.
Australia's wobbegong sharks are flattened ambush hunters fringed with weed-like skin flaps, camouflaged so well against reef floors that their name is thought to come from an Aboriginal word associated with shaggy beards. Prey swims to them. The rug moves last.
In round-the-clock observations at 22 °C (72 °F), nocturnal octopuses averaged about 12 quiet-sleep bouts of 50 minutes each plus roughly 10 brief active-sleep bouts. Together, these amounted to about 10 hours of sleep per day. Warmer water shortened the interval between active bouts.
The octopus genome contains 168 protocadherin genes, which help regulate how neurons develop and connect. That is about ten times the number found in other invertebrates and more than twice the number in mammals. Large protocadherin families had previously been considered mainly a vertebrate feature.
The silky shark's denticles are so small and densely packed that its skin feels sleek rather than sandpapery — the trait that named it. The finish trims drag for a life spent chasing tuna in open water. Even among sharks, there is a best-dressed.
The shark lineage has endured for more than 400 million years, swimming through all five of Earth's great mass extinctions — including the one that ended the dinosaurs. They watched the rise and fall of nearly everything else in the ocean.
Vision-model research simulating a white shark's eye found that surfers paddling overhead closely match the silhouette and motion of seals — supporting the long-held mistaken-identity explanation for many bites, which are frequently single strikes without feeding. To a young white shark looking up, a surfboard is a seal-shaped question.
The whale shark is the largest fish alive, and the biggest reliably measured individual reached 18.8 m (61 ft 8 in). It feeds almost entirely on plankton, straining tiny drifting life through a mouth up to 1.5 m (5 ft) wide. The ocean's biggest shark eats some of its smallest creatures.
Scientists fitted tiger sharks in the Bahamas with cameras and trackers, and the sharks' commutes across their feeding grounds helped confirm the largest known seagrass ecosystem on Earth — at least 66,000 km² (25,000 sq mi) and up to 92,000 km² (35,000 sq mi) across the Bahama Banks, enough to raise the world's known seagrass cover by more than 40% in a 2022 study. The apex predator moonlighted as a survey vessel.
Juvenile lemon sharks in Bimini prefer specific companions and learn tasks faster after watching experienced sharks — evidence of social personality and social learning in a fish long assumed solitary and simple. Shark school is not just a pun.
Small deep-sea lantern sharks grow light-producing organs called photophores along their undersides, matching the dim glow filtering from above so predators looking up see nothing. Some also flash light along spines and fins. In the deep, the best hiding place is brightness.
Because cartilage rarely fossilizes, the shark fossil record is overwhelmingly teeth — shed continuously for 400 million years — plus occasional vertebrae and rare body impressions. Some fossil beds hold thousands of teeth per square meter. Sharks wrote their autobiography almost entirely in bite.
Every whale shark wears a unique constellation of pale spots, so scientists adapted a 1986 star-mapping algorithm built for the Hubble Space Telescope era to match shark flanks the way astronomers match star fields. The method anchors a global photo-ID library with tens of thousands of encounters. Fittingly, the whale shark's name in Madagascar — marokintana — means "many stars."
Radiocarbon from Cold War bomb tests, lodged in the growth bands of two dead whale sharks' vertebrae, proved in 2020 that each band marks one year — and put one shark, a 10 m (33 ft) female stranded in Pakistan, at 50 years old, the oldest ever verified. Earlier modelling built on those yearly bands suggests the largest may live a century. The biggest fish in the sea is also one of its slow-burning elders.
A shark's skin is armored with tooth-like denticles whose ridges shed drag, and engineers have been copying them. In 2014 a Harvard team 3D-printed an artificial shark skin and showed it improved swimming performance on a flapping foil; in 2018 the same lab, with the University of South Carolina, molded shortfin mako denticles onto an airfoil and found arrangements that cut drag and added lift at once, improving the lift-to-drag ratio by up to 323% at low angles of attack. Denticle-patterned surfaces are studied for aircraft, drones and wind turbines. Four hundred million years of prototyping ended up in engineering journals.
Feeding trials published in 2018 showed bonnethead sharks digesting and absorbing nutrients from seagrass, not merely swallowing it while hunting crabs — the first shark shown to be a true omnivore. More than half its diet can be plants. Somewhere in the hammerhead family, a shark went partly vegetarian.
In April 2019, cameras lowered into the Java Trench during the Five Deeps Expedition recorded a dumbo octopus foraging at 6,957 meters (22,825 feet), in water of about 1.5 °C (35 °F). A second animal turned up at 5,760 meters (18,898 feet). The previous reliable record, a black-and-white photograph, had stood at 5,145 meters for half a century.
The white shark's two-tone paint job is engineering: slate-gray above blends into deep water for eyes looking down, while the pale belly dissolves into surface glare for eyes looking up. The scheme, countershading, hides a two-ton predator in open water. The most famous silhouette in the sea is built to have none.
The cookiecutter's lower teeth are fused into a single serrated band that sheds whole, not tooth by tooth — and the shark swallows the discarded band, likely recycling its calcium. It eats pieces of giants, and then it eats its own knives. Nothing in its economy goes to waste.
In History of Animals, Aristotle called the octopus a stupid creature because it would approach a person's lowered hand. Yet he also accurately described its specialized mating arm, a feature European naturalists later had to rediscover.
Radiocarbon dating of proteins in their eye lenses shows Greenland sharks live at least 272 years — longer than any other animal with a backbone. The oldest of the 28 sharks measured in the landmark 2016 study, a female just over 5 m (16 ft) long, was estimated at roughly 392 years old. Females don't even reach maturity until around 150, and a big one cruising the Arctic today has been swimming for some three centuries.
Sharks have no gas-filled swim bladder; buoyancy comes from an enormous oil-rich liver that can approach a quarter of a shark's weight in some species. The oil, squalene, is lighter than water, letting a heavy cartilage body hang almost weightless. A shark floats on its own chemistry.
Voluntary arm shedding, or autotomy, has been documented in at least Abdopus aculeatus and Ameloctopus litoralis. The arm separates at a defined break point and continues writhing while the octopus escapes.
The great white shark gives more of its brain to smell than any other shark: its olfactory bulbs — the paired lobes that process scent — run to about 18% of its brain mass.
Octopuses drag prey home and discard the shells outside the entrance, building a heap called a midden. Divers use these piles of cracked crab and clam shells to locate dens that would otherwise be invisible. The habit is old enough that Aristotle noted it more than 2,000 years ago.
Male and juvenile blanket octopuses tear stinging tentacles from Portuguese men-of-war and hold them in their arms, apparently as weapons or deterrents. Skin chemistry is thought to keep the stinging cells from firing against them.
The thresher's scythe-like upper tail can equal the length of its body, and high-speed video published in 2013 shows the shark whipping it overhead to stun whole groups of sardines before circling back to swallow the dazed. It is the only shark that hunts with a bullwhip.
The International Shark Attack File puts a U.S. beachgoer's lifetime-scale risk of a fatal shark bite near one in 4.3 million — far below the risks of lightning, bees or the drive to the beach. In a typical year sharks bite about 70 people worldwide, unprovoked, and about six of those bites are fatal.
Pairs of larger Pacific striped octopuses mate face to face with their suckers aligned, share food and may occupy the same den for days. Females produce multiple clutches over months instead of a single terminal brood.
Shark jaws hang in slings of muscle and ligament, letting the whole apparatus thrust forward and gape wide independent of the head — the mechanism behind slow-motion footage of white sharks seeming to launch their mouths at prey. The skull aims; the jaw travels.
Two branchial hearts drive blood through the gills; a third systemic heart sends it out to the arms, organs and brain. When the animal jets, pressure inside the mantle stops blood returning, and the systemic heart — with nothing to pump — stops until the jetting ends. Fast swimming therefore starves an octopus of oxygen at the exact moment it needs it most. This makes sustained jetting physiologically costly compared with crawling.
The hammerhead's wide-set eyes give it a panoramic view and, when it sweeps its head, effectively 360-degree vision — a 2009 study also confirmed excellent stereo depth perception from the exaggerated eye separation. The strangest head in the sea turns out to be a surveying instrument.
A 2009 genetic survey found venom proteins across octopuses, cuttlefish and some squid, tracing them to a single ancient venomous ancestor. Most species use theirs quietly, paralyzing a clam until its shell relaxes open. Only the blue-ringed octopuses of the genus Hapalochlaena carry a toxin dangerous to humans.
A whale shark's back is covered in some of the thickest skin in the animal kingdom — up to about 10 cm (4 in) — and it hardens the surface further by flexing when boats or predators come close. The gentlest giant wears the heaviest coat.
On Great Barrier Reef platforms cut off at night by spring low tides, the epaulette shark tolerates hours of severe hypoxia — oxygen near 5% of normal — and at least an hour of total anoxia, with no drop in its brain's energy stores and no neurons lost afterward. It manages this by throttling demand rather than boosting supply: an adenosine-driven slowdown leaves the shark in a coma-like torpor until oxygen returns, when it snaps back to alertness within moments. Researchers at Griffith University and the University of Auckland study its brain mitochondria for clues to limiting stroke and heart-attack damage, where the harm comes as oxygen floods back. A reef shark's hour without oxygen is a lesson medicine wants.
By 2020, pocket sharks were known from exactly two specimens: a 40 cm (16 in) female trawled near the Nazca Ridge off Chile in 1979, and a 14 cm (5.5 in) male netted in the Gulf of Mexico in 2010 and named the American pocket shark in 2019. Each carries a slit-like pocket above the base of each pectoral fin, and a 2020 study of the Gulf specimen's pocket glands found the cellular machinery for secreting a luminous fluid. A shark barely longer than a smartphone is built with glowing pockets.
Juvenile scalloped hammerheads cruising sun-drenched shallows darken dramatically as melanin builds in their skin — one of the first demonstrations of true sunlight-driven tanning in a fish, published in Nature in 1996. Sharks kept shaded stayed pale while sunlit sharks browned. The beach tan predates the beach towel by millions of years.
Portuguese dogfish have been recorded around 3,700 m (12,100 ft) deep — among the deepest confirmed for any shark — hunting in freezing blackness at pressures hundreds of times the surface's. Even the abyss has its patrols.
The star-sucker pygmy octopus, Octopus wolfi, lives in the shallow coastal waters of the western Pacific Ocean, measures under 2.5 centimeters (1 inch) in total length and weighs less than a gram (0.04 ounce) — smaller and lighter than a small coin, and small enough to sit on a fingernail. It is named for the ring of tiny papillae fringing the suckers near its arm tips, which let it taste whatever it touches.
When an octopus cannot pull a shell open, a muscular salivary papilla studded with minute teeth — separate from the radula, its primary toothed ribbon — can drill through it. Secretions from the posterior salivary glands then help relax the muscle holding the shell closed.
The bluntnose sixgill belongs to the Hexanchiformes, the oldest-known group of modern sharks, already swimming by the Early Jurassic about 195 million years ago — before most dinosaurs, though not before the first. It keeps six gill slits where nearly every other living shark has five, and patrols deep water worldwide, rising toward the surface at night. Meeting one is the closest thing to diving with the Jurassic.
Greenland shark meat is loaded with trimethylamine oxide and urea, enough to sicken anyone who eats it fresh. Icelanders ferment and air-dry it for months into hákarl, a pungent national delicacy served in small cubes. The world's longest-lived vertebrate is also one of its most acquired tastes.
The bold dark bars that name the tiger shark belong mostly to youth: they fade as the shark matures, leaving big adults far plainer than their portrait. The name records the juvenile. By the time a tiger shark is truly formidable, it has nearly erased its tiger.
During more than 500 diver hours off Indonesia, researchers watched veined octopuses dig up coconut-shell halves, stack them and carry them as far as 20 meters (66 feet) before assembling them into shelters. Because the shells provide no protection while being transported and are saved for later use, the behavior was classified as tool use.
Chromatophores hold only yellow, red, brown and black pigments, so no combination of them can produce the blue-green sheen a living octopus sometimes shows. Two other cell types do that work: iridophores, stacks of protein plates that split light by interference, and leucophores, which scatter whatever light falls on them and so look red in red light and blue in blue. Part of octopus color is not paint at all. It is optics.
That is every fact on the shelf — all 199 of them.
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